研究生: |
姚愛文 Ai- Wen Yao |
---|---|
論文名稱: |
福山試驗林孔隙特性與孔隙-非孔隙林下苗木組成之研究 Gap characteristics and seedling composition within gap and non-gap understory of Fushan Experimental Forest |
指導教授: |
林登秋
Lin, Teng-Chiu |
學位類別: |
碩士 Master |
系所名稱: |
生命科學系 Department of Life Science |
論文出版年: | 2013 |
畢業學年度: | 101 |
語文別: | 中文 |
論文頁數: | 69 |
中文關鍵詞: | 孔隙特性 、福山試驗林 、苗木組成 、颱風 、林下光 |
英文關鍵詞: | gap characteristics, Fushan Experimental Forest, seedling composition, typhoon, understory light |
論文種類: | 學術論文 |
相關次數: | 點閱:75 下載:9 |
分享至: |
查詢本校圖書館目錄 查詢臺灣博碩士論文知識加值系統 勘誤回報 |
林冠孔隙的出現提供額外的空間和光等資源,讓林下耐陰樹種有機會加速生長向林冠邁進,因此孔隙可促進冠層樹種的更新,並維持森林物種多樣性。熱帶與溫帶地區,已有大量關於孔隙的研究,但卻少有研究在亞熱帶森林進行。台灣位處亞熱帶,又位於北太平洋西岸,颱風侵襲最頻繁的地區,使森林因應颱風擾動而產生結構上的適應。本研究的目的即在探討福山,一個經常受颱風擾動的亞熱帶森林,其孔隙的分布和特性與其它地方對孔隙的研究結果是否有所不同;如頻繁的颱風使樹木定期落葉,提供林下相對較高的光量,可能使耐陰樹種的小苗無需依賴孔隙即能向上生長至林冠,因此森林是否不需依賴孔隙進行更新?本研究在福山試驗林一號試驗集水區設置七條由稜線延伸至溪谷的樣線,調查所經過的孔隙,內容包含孔隙面積、年齡,及計算森林的更新時間(turnover time),並與1995年前人在1994年的六個颱風過後,於福山進行的孔隙研究做一比對。研究中亦調查苗木組成及測量林下光,以比較孔隙和非孔隙林下的苗木組成與光環境的關係。研究共記錄17個孔隙,孔隙面積在1-36 m2之間,數量和面積皆較過去的研究為少和小。林份更新時間以形成1年與9年內的孔隙做計算分別為197年和398年,遠較過去的研究所估算的51年和175年為長。林下光照在孔隙和非孔隙間沒有顯著差異,此結果反應在林下苗木於孔隙和非孔隙的組成相似性高,且非孔隙林下擁有孔隙中所有出現的苗木物種,顯示颱風擾動頻繁的福山林木似不需依賴孔隙進行更新。
Forest gaps can promote canopy tree regeneration thereby maintain forest species diversity. Gaps provide space and light for rapid growth of understory plants, especially shade-tolerant species, thus help them reach the canopy. This regeneration process has been well described in tropical and temperate forests, but is rarely studied in subtropical forests. Forests in Taiwan which is located at the west side of North Pacific, is influenced by frequent typhoons as such the forests have structural adaptation to typhoon disturbance. This study examined gap characteristics in a subtropical forest of northeastern Taiwan, the Fushan Experimental Forest. Frequent typhoon disturbance causes regular defoliation, and thereby increases understory light availability allowing seedlings of shade-tolerant species to reach the canopy. It is questionable that in such a forest, if gap dynamics also plays an important role on canopy tree regeneration. To address this question, 7 transects were set up at watershed #1 in Fushan Experimental Forest. The gaps intersected the transect lines were surveyed and gap characteristics including area size, age, and turnover time were compared to the study conducted in 1995 following the disturbance of 6 typhoons of 1994. Seedling composition and understory light environment were also investigated to examine their relation to gap and non-gap understory. A total 17 gaps were recorded and the area of individual gaps ranged from 1 to 36 m2. Both the numbers and size of the gaps were less/smaller than those reported before. The calculated turnover time is 197 and 398 years when gaps < 1 and ≦ 9 years, respectively, were include and is much longer than the 51 and 175 years reported in the 1995 study. Understory light availability showed no significant difference between gap and non-gap understory and this contributed to the high similarity of seedling composition between the two environments. Moreover, non-gap understory encompasses all seedling species recorded in gaps. Thus, seedling regeneration in Fushan may not rely on gaps.
中央氣象局。2013。中央氣象局TDB防災颱風資料庫網頁系統,http://rdc28.cwb.gov.tw/data.php於 2013 年 5 月2日瀏覽。
王相華、潘富俊、劉景國、于幼新和洪聖峰。2000。台灣北部福山試驗林永久樣區之植物社會分類及梯度分析。台灣林業科學15 (3):411-428。
林元祥。2007。福山天闊葉林內光環境對種苗存活與生長表現之影響。國立東華大學自然資源管理研究所碩士論文。
林文雄。2008。福山試驗林幼木生長與立地條件因子關係之探討。國立彰化師範大學地理學系碩士論文。
林登秋和江智民。2002。半球面影像在森林生態研究的應用。台灣林業科學17 (3):387-400。
夏禹九和黃正良。1999。福山試驗林的水文特性。中華林學季刊 32 (1):39-51。
夏禹九、黃正良和陳明杰。1996。福山試驗集水區量水堰之設計-分段複式量水堰。台灣林業科學11 (2):113-122。
陳明義、周文郅、蔡進來。2000。關刀溪森林生態系之倒木孔隙更新。林業研究季刊22 (1):23-32。
康若蘭、林登秋、詹進發和黃正良。2000年碧利斯颱風干擾前後福山試驗林NDVI變動之研究。台灣林業科學20 (1)::73-87。
曾維宏。1994。南仁山區低海拔亞熱帶雨林林隙更新之研究。國立臺灣大學植物學研究所碩士論文。
張乃航、馬復京、游漢明和許原瑞。1998。福山地區次生闊葉林土壤種子庫及幼苗動態。台灣林業科學13 (4):279-289。
劉棠瑞和蘇鴻傑。1983。森林植物生態學。台北:台灣商務印書館股份有限公司。頁:9。
Abe, S., Masaki, T., Nakashizuka, T. 1995. Factors influencing sapling composition in canopy gaps of a temperate deciduous forest. Vegetatio 120: 21-32.
Barden, L.S. 1980. Tree Replacement in a Cove Hardwood Forest of the Southern Appalachians. Oikos 35: 16-19.
Bray, J. R. & Curtis, J.T. 1957. An ordination of the upland forest communities of southern Wisconsin. Ecological Monograph 27: 325-349.
Bray, R.J. 1956. Gap phase replacement in a Maple-Basswood forest. Ecology 37: 598-600.
Brokaw, N.V.L. 1982. The definition of treefall gap and its effect on measures of forest dynamics. Biotropica 14:158-160.
Brokaw, N.V.L. 1985a. Gap-phase regeneration in a tropical forest. Ecology 66: 682-687.
Brokaw, N.V.L. 1985b. Treefalls, regrowth, and community structure in tropical forest. In: Pickett, S.A., White, D.C. (Eds.), The Ecology of Natural Disturbance and Patch Dynamics. pp. 53-69. Academic Press, New York.
Canham, C.D., Denslow, J.A., Platt, W.J., Runkle, J.R., Spies, T.A., and White, P.S. 1990. Light regimes beneath closed canopies and tree-fall gaps in temperate and tropical forests. Canadian Journal of Forest Research 20: 620-631.
Chazdon, R.L., and Fetcher, N. 1984. Photographic estimation of photosynthetically active radiation: evaluation of a computerized technique. Oecologia 72:553-564.
Clark, D.B., Clark, D.A., Rich, P.M., Weiss, S.B., and Oberbauer, S.F. 1996. Landscape-scale evaluation of understory light and canopy structure: methods and application in a neotropical lowland rain forest. Canadian Journal of Forest Research 26:747-757.
Clebsch, E.E.C. and Busing, R.T. 1989. Secondary succession, gap dynamics, and community structure in a southern appalachian cove forest. Ecology, 70(3): 728-735.
Cooper, W.S. 1913. The climax forest of Isle Royale, Lake Superior, and its development. Botanical Gazette 55: 1-44.
Denslow, J.S. 1987. Tropical rain-forest gaps and tree species-diversity. Annual Review of Ecology and Systematics 18: 431-451.
Fraver, S., Brokaw, N.V.L., and Smith, A.P. 1998. Delimiting the gap phase in the growth cycle of a Panamanian forest. Journal of Tropical Ecology 14: 673-681.
Gray, A.N., Spies, T.A., and Easter, M.J. 2002. Microclimatic and soil moisture responses to gap formation in coastal Douglas-fir forests. Canadian Journal of Forest 32: 332-343.
Hill, M.O. 1973. Diversity and evenness: a unifying notation and its consequences. Ecology 54: 427-431.
Krasny, M.E., Whitmore, M.C. 1992. Gradual and sudden forest canopy gaps in Allegheny northern hardwood forests. Canadian Journal of Forest Research 22: 139-143.
Lieberman, D., Liebermarn, M., Peralta, R., and Hartshorn, G.S. 1985. Mortality patterns and stand turnover rates in a wet tropical forest in Costa Rica. Ecology 73: 915-924.
Lin, T.C., Hamburg, S.P., Hsia, Y.J., Lin, T.T., King, H.B., Wang, L.J., and Lin, K.C. 2003. Influence of typhoon disturbances on the understory light regime and stand dynamics of a subtropical rain forest in northeastern Taiwan. Journal of Forest Research 8:139-145.
Lin, T.C., Hamburg, S.P., Lin, K.C., Wang, L.J., Chang, C.T., Hsia, Y.J., Vadeboncoeur, M.A., Mabry McMullen, C.M. and Liu, C.P. 2011. Typhoon disturbance and forest dynamics: Lessons from a Northwest Pacific subtropical forest.
Ecosystems 14: 127-143.
Mabry, C.M., Hamburg, S.P., Lin, T.C., Horng, F.W., King, H.B. and Hsia, Y.J. 1998. Typhoon disturbance and stand-level damage patterns at a subtropical forest in Taiwan.
Biotropica 30(2): 238-250.
McCarthy, J. 2001. Gap dynamics of forest trees: A review with particular attention to boreal forests. Environmental Reviews 9(1): 1–59.
McNab, W.H., Greenberg, C.H., and Berg, E.C. 2004. Landscape distribution and characteristics of large hurricane-related canopy gaps in a southern Appalachian watershed. Forest Ecology and Management 196: 435-447.
Muscolo, A., Sidari, M., and Mercurio, R., 2007. Influence of gap size on organic matter decomposition, microbial biomass and nutrient cycle in Calabrian pine (Pinus laricio, Poiret) stands. Forest Ecology and Management 242: 412-418.
R Core Team 2013. R: A language and environment for statistical computing. R Foundation for Statistical Computing, Vienna, Austria. ISBN 3-900051-07-0, URL http://www.R-project.org/.
Robin, D., Lesage, I., Messier, C., and Morin, H. 2001. Effects of light and intraspecific competition on growth and crown morphology of two size classes of understory balsam fir saplings. Forest Ecology and Management 140: 215-225.
Runkle, J.R. 1981. Gap regeneration in some old-growth forests of the Eastern United-States. Ecology 62: 1041-1051.
Runkle, J.R. 1982. Patterns of disturbance in some old-growth mesic forests of eastern north america. Ecology 63(5): 1533-1546.
Runkle, J.R. 1985. Disturbance regimes in temperate forests. In: Pickett, S.A., White, D.C. (Eds.), The Ecology of Natural Disturbance and Patch Dynamics. pp. 17-33. Academic Press, New York.
Runkle, J.R. 1992. Guidelines and sample protocol for sampling forest gaps. USDA Forest Service Pacific Northwest Research Station General Technical Report PNW-GTR-283. pp. 44.
Schliemann, S.A., and Bockheim, J.G. 2011. Methods for studying treefall gaps: A review. Forest Ecology and Management 261: 1143-1151.
Schnitzer, S.A., and Carson, W.P. 2001. Treefall gaps and the maintenance of species diversity in a tropical forest. Ecology 82: 913-919.
Schnitzer, S.A., Mascaro, J., Carson, W.P. 2008. Treefall gaps and the maintenance of species diversity in tropical forest. In: Carson, W.P., Schnitzer, S.A. (Eds), Tropical Forest Community Ecology. pp196-209. Wiley-Blackwell, Oxford, UK.
Turton, S.M. 1992. Understory light environments in a northeast Australian rain forest before and after a tropical cyclone. Journal of Tropical Ecology 8: 241-252.
Uhl, C., Clark, K., Dezzeo, N., and Maquino, P. 1988. Vegetation dynamics in Amazonian treefall gaps. Ecology 69: 751-763.
Watt, A.S., 1947. Pattern and process in the plant community. Journal of Ecology 35: 1-22.
White, P.S., 1979. Pattern, process, and natural disturbance in vegetation. Botanical Review 45: 229-299.
White, P.S. and Pickett, S.T.A. 1985. Natural disturbance and patch dynamics: an introduction. In: Pickett, S.A., White, D.C. (Eds.), The Ecology of Natural Disturbance and Patch Dynamics. pp. 3-9. Academic Press, New York.
Whitmore, T.C. 1989. Canopy gaps and the two major groups of forest trees. Ecology 70: 536-538.
Williams, H., Messier, C., and Kneeshaw, D. 1999. Effects of light availability and sapling size on the growth and crown morphology and understory Douglas-fir and lodgepole pine. Canadian Journal of Forest Research 29: 222-231.
Yamamoto, S.I. 1992. The gap theory in forest dynamics. Botanical Magazine (Tokyo) 105: 375-383.
Yamamoto, S.I. 2000. Forest gap dynamics and tree regeneration. Journal of Forest Research 5: 223-229.
Zhang, Q.H., and Zak, J.C., 1995. Effects of gap size on litter decomposition and microbial activity in a subtropical forest. Ecology 76: 2196-2204.