動物的移動能力經常受到環境因子與本身的形態結構影響,而適應於一種類型環境的形態結構也經常會影響動物在␣一類型環境 時的運動表現。
闊尾海蛇(Laticaudinae)在生活史上和陸地保有一定程度的關連性, 並且因種間水棲化程度的不同,在形態與生理上也有不同程度的特 化。本實驗預期:水棲化程度最強的闊帶青斑海蛇(Laticauda semifasciata)最偏好在水域環境,且有最弱的陸地移動表現,並反應 在運動形態、攀爬、水面抬升、跨越能力以及肌肉拉力的表現上; ␣唇青斑海蛇(L. colubrina)則最偏好陸域環境,且有最強的陸地移動 表現;黑唇青斑海蛇(L. laticaudata)則介於兩者之間。結果顯示三種 闊尾海蛇一天當中在陸地上的時間比例沒有顯著差異,且黑唇青斑 海蛇的水陸交換次數最多。在運動形態方面三種闊尾海蛇均表現出 不會利用支點移動的傾向,然而陸棲的近␣物種雨傘節卻可利用支點做蜿蜒爬行的動作,可能是 因為演化上闊尾海蛇的親␣關係較接近於真海蛇,導致在行為上與 陸棲蛇類之間有所差異。而在陸地移動能力方面,闊帶青斑海蛇經 常有最好的表現,黑唇青斑海蛇則最差,可能是由於三種物種本身 體形上的差異與肌肉含量的不同,造成不合乎水棲化程度影響陸地 移動表現的預期。而由於陸地的移動能力在闊尾海蛇的生活史上有 時會直接影響到生殖成功的機會,因此若是以與吻肛長的比例表示 其陸地移動能力時,三種闊尾海蛇之間則沒有顯著差異。

The locomotion abilities of animals are usually affected by environmental factor and the morphologies of its self, and the structure that adaptive to one kind of environment may affect the locomotion expression in other kind of environment.
The life history of Laticaudinae maintains a certain level of relationship with the land, and carry different levels of morphological and physical specialization caused by different aquatic levels between species. The study predict that: Laticauda semifasciata, the most aquatic species, will prefer the aquatic environment, and have the weakest locomotor abilities on land, reflect in moving performance, climbing ability, raising from water, and muscle strength; And the most terrestrial species, L. colubrine, prefer the terrestrial environment, and have the strongest terrestrial locomotor abilities. And the locomotor abilities of L. laticaudata are between the L. semifasciata and L. colubrine. The result show that the L. colubrine stay the longest time period on land. The three kinds of laticauda sea snakes show the same moving performance without using the pivot points, and totally different with the terrestrial species Bungarus multicinctus multicinctus. Its may because of that Laticaudinaes are much closer with the true sea snakes. In the aspect of terrestrial locomotor abilities, L. semifasciata and L. colubrina usually have the same represent, and also usually better than L. laticaudata. The reason of results unmatching the predictions might be caused by the differences of the morphology and muscle contains between these three
species.

Ashley-Ross, M.A., and Bechtel, B.F. 2004. Kinematics of the transition between aquatic and terrestrial locomotion in the newt Taricha torosa. J. Exp. Biol.(207):461-474.
Barbosa, A., and Moreno, E. 1999. Hindlimb morphology and locomotor performance in waders: an evolutionary approach. Biol. J. Linn. Soc.(67): 313-330.
Bennett, A.F., Losos, J.B., and Walton, B.M. 1993. Trade-offs between sprinting and clinging ability in Kenyan chameleons. Funct. Ecol.(7):281-286.
Biewener, A.A., and Corning, W.R. 2001. Dynamics of mallard(Anas platyrhyncos) gastrocnemius during swimming versus terrestrial locomotion. J. Exp. Biol.(204):1745-1756.
Blob, R.W., Bridges, W.C., Ptacek, M.B., Maie, T., Cediel, R.A., Bertolas, M.M., Julius, M.L., and Schoenfuss, H.L. 2008.
Morphological selection in an extreme flow environment: body shape and waterfall-climbing success in the Hawaiian stream fish Sicyopterus stimpsoni. Integr. Comp. Biol.(48)6:734-749.
Bogert, C.M. 1947. Rectilinear locomotion in snakes. Copeia(1947):253-254.
Bonnet, X., Ineich, I., and Shine, R. 2005. Terrestrial locomotion in sea snakes: the effects of sex and species on cliff-climbing ability in sea kraits (Serpentes, Elapidae, Laticauda). Biol. J. Linn. Soc.(85):433-441.
Bonnet, X., Shine, R., Naulleau, G., Vacher-Vallas, M. 1998. Sexual dimorphism in snakes: different reproduction roles favour different body plans. Proc. R. Soc. Lond. B. Biol. Sci.(265):1-5.
Bradshaw, S.D., Gans, C., and Saint-Girons, H. 1980. Behavioural thermoregulation in a pygopodid lizard, Lialis burtonis. Copeia.(980):7380-743.
Brischoux, F., Bonnet, X., and Pinaud, D. 2009. Fine scale site fidelity in sea kraits: implications for conservation. Biodivers Conserv.(18):2473-2481.
Buffa, P. 1904. Ricerche Sulla Muscolatura Cutanea Dei Serpenti e Considerazioni sulla di questi animali. Dall’l stituto di Zoologia e Amatomia comparata della R. Unicersita di Padova. pp.89.
Cadle, J.E., and Gorman, G.C. 1981. Albumin immunological evidence and the relationships of sea snakes. J. Herpetol.(15):329-334.
Cartmill, M. 1985. Climbing. In: “Functional Vertebrate Morphology” ed. by M. Hildebrand, D.M. Bramble, K.F. Liem, and D.B. Wake(eds.), pp.73-88. The Belknap Press, Cambridge.
Cundall, D. 1987. Functional Morphology. In “Snake: Ecology and Evolutionary Biology” ed. by R.A. Seigel, J.T. Collins, and S.S. Novak(eds.), Collier Macmillan Canada, Inc. New York, pp106-140.
Damme, Van R., Aerts, P., and Vanhooydonck, B. 1997. No trade-off between sprinting and climbing in two populations of the lizard podarcis hispanica(Reptilia: Lacertidae). Biol. J. Linn. Soc.(60):493-503.
Dobson, F.S. 1992. Body mass, structural size, and life- history patterns of the Columbian ground squirrel. Am. Nat.(140):109-125.
Domonici, P., and Blake, R.W. 1991. The kinematics and performance of the escape response in the angelfish(Pterophyllum eimekei). J. Exp. Biol.(156):187-205.
Dullemeijer, P. 1974. Concepts and Approaches in Animal Morphology. Van Gorcum, Assen, The Netherlands.
Gans, C. and Gasc, J.P. 1990. Tests on the locomotion of the elongate and limbless reptile Ophisaurus apodus(Sauria: Anguidae). J. Zool.(220):517-536.
Gillis, B.G. 1996. Undulatory locomotion in elongate aquatic vertebrates: anguilliform swimming since Sir James Gray. Amer. Zool.(36):656-665.
Gillis, B.G. 1998. Environmental effects on undulatory locomotion in the American eel(Anguilla rostrata): kinematics in water and land. J. Exp. Biol.(201):949-961.
Graham, J.B., and Lowell, W.R. 1987. surface and subface swimming of the sea snake Pelamis Platurus. J. Exp. Biol.(127):27-44.
Gray, J. 1953.Undulatory propulsion. Quart. J. Micro. Sci.(94):551-578.
Gray, J., and Lissmann, H.W. 1950. The kinetics of locomotion of the grass snake. J. Exp. Biol.(26):354-367.
Guinea, M.L. 1986. Aspects of the biology of the common Fijian sea snake. Laticauda Colubrina(Schneider). University of the South Pacific, Suva, Fiji.
Guyer, C., and Donelly, M.A. 1990. Length mass relationship among as assemblage of tropical snakes in Costa Rica. J. Trop. Ecol.(6):65-76.
Harding, K.A., and Welch, K.R.G. 1908. Venomous Snakes of the World, A Checklist. Pergamon Press, New York.
Heatwole, H. 1999. Sea Snakes. University of NSW Press, Sydney, NSW.
Hirth, H.F., and King, A.C. 1969. Body temperatures of snakes in different seasons. J. Herpetol.(3):101-102.
Ineich and Laboute. 2002. Les Serpents Marines de Nouvelle-Calidonie/Sea Snakes of New Caledonia. Institut de Recherche pour le Developpement, Museum national d’Histoire naturelle, Paris.
Irschick, D.J. 2003. Studying performance in nature: implications for Fitness variation within populations. Integrative and Comparative Biology(43):396-407.
Jayne, B.C. 1982. Comparative morphology of the semispinalisspinalis muscle of snakes and correlations with locomotion and constriction. J. Morphol.(172): 83-96.
Jayne, B.C. 1988a. Muscular mechanisms of snake locomotion: an Electromygraphic study of lateral undulation of the Florida banded water snake(Nerodia fasciata) and the yellow rat snake(Elaphe obsoleta). J. Morphol.(197):159-181.
Jayne, B.C. 1988b. Muscular mechanisms of snake locomotion: an electromyographic study of the sidewinding and concertina modes of Crotalus cerastes, Nerodia fasciata and Elaphe obsoleta. J. Exp. Biol.(140):1-33.
Jayne, B.C., and Raley, M.C. 2007. Scaling of the axial morphology and gap-bridging ability of the brown tree snake(Boiga irregularis). J. Exp. Biol.(210):1148-1160.
Keogh, J.S. 1997. ‘‘Phylogenetic Studies on Australo-Papuan Elapid Snakes Based on Morphology and DNA,’’ Unpublished Ph.D Thesis, University of Sydney, Australia.
Keogh, J.S. 1998. Molecular phylogeny of elapid snakes and a consideration of their biogeographic history. Biol. J. Linn. Soc.(63):177-203.
Kitchell, J.F. 1969. Thermophilic and thermophobic responses of snakes in a thermal gradient. Copeia(1969):189-91.
Lillywhite, H.B. 1985. Behavioral control of arterial pressure in snakes. Physiol. Zool.(58)159-165.
Lillywhite, H.B., and Henderson, R.W. 1993. Behavioral and function ecology of arboreal snakes. In “Snakes: Ecology and Behavior” ed. by R.A. Seigel and J.T. Collins(eds.), MacGraw-Hill, Inc. New York, pp.1-48.
Lillywhite, H.B. 1987. Circulatory adaptations of snakes to gravity. Am. Zool.(27):81-95.
Lillywhite, H.B. 1987. Temperature, energetics, and physiological ecology. In “Snakes: Ecology and Evolutionary Biology.”ed. by R.A. Lillywhite, H.B., Babonis, L.S., Sheehy, C.M. 3rd., and Tu, M.C. 2008. Skin permeability and water relations of sea snakes. FASEB J.(22):1239.18.
Lillywhite, HB., LaFrentz, J.R., Lin, Y.C., and Tu, M.C. 2000. The cantilever abilities of snakes. J. Herpetol.(3):523-528.
Lillywhite, H.B., Zipple, K.C., and Farrel, A.P. 1999. Resting and maximal heart rates in ectothermic vertebrates. Comp. Biochem. Physiol.(124)A:369-384.
Losos, J.B., Irschick, D.J., and Schoener, T. 1994. Adaptation constraints in the evolution of specialization of Bahamian Anolis lizards. Evolution(48): 1786-1798.
Losos, J.B. and Sinervo, B. 1989. The effects of morphology and perch diameter on sprint performance of Anolis lizards. J. Exp. Biol.(245):23-30.
Losos, J.B., Walton B.M., and Bennett, A.F. 1993. Trade-offs between sprinting and climbing ability in Kenyan chameleons. Funct Ecol.(7):281-286.
Lourdais, O., Brischoux, F., De Nardo, D., and Shine, R. 2004. Protein catabolism in pregnant snakes(Epicrates cenchria maurus Boidae) compromises musculature and performance after reproduction. J. Comp. Physiol. B. Biochem. Syst. Environ. Physiol.(174):383-391.
Lutterschmidt, W.I., and Reinert, H.K. 1990. The effect of ingested transmitters upon the temperature preference of the northern water snake, Nerodia s. sipedon. Herpetologica(46):39-42.
Mao, S.H., Chen, B.Y., and Chang, H.M., 1977. The evolutionary relationships of sea snakes suggested by immunological cross-reactivity of transferrins. Comp. Biochem. Physiol.(57A)4: 403-406.
Mao, S.H., Chen, B.Y., Yin, F.Y., and Guo, Y.W. 1983. Immunotaxonomic relationships of sea snakes to terrestrial elapids. Comp. Biochem. Physiol.(74A)4:869-872.
McDowell, S.B. 1972. The genera of sea-snakes of the Hydrophis group(Serpentes, Elapidae). Trans. Zool. Soc. Lond.(32):189-247.
Mosauer, W. 1932. On the locomotion of snakes. Science(76):583-585.
Mullin, J.S. and Cooper, J.R. 2002. Barking up the wrong tree: climbing performance of rat snakes and its implications for depredation of avian nests. Can. J. Zool.(80): 591-595.
Naulleau, G. 1979. Etude biotélémétrique de la thermoregulation chez Vipera aspis(L.) (Reptilia, Serpentes, Viperidae) elévee en conditions artificielles. J. Herpetol.(13):203-208.
Pfennig, D.W., Rice, A.M., and Martin, R.A. 2007. Field and experimental evidence for competition’s role in phenotype divergence. Evolution(61)2:257-271.
Pernetta, J.C. 1977. Observations on the habits and morphology of the sea snake Laticauda colubrina(Schneider) in Fiji. Can. J. Zool.(55):1612-1619.
Pounds, J.A. 1988. Ecomorphology, locomotion, and microhabitat structure: patterns in a tropical mainland Anolis community. Ecol. Monogr.(58):299-320.
Pough, F.H., Andrews, M.B., Cadle, E.J., Crump, L.M., Savitzky, H.A., and Wells, D.K. 1998. Herpetology. Premtice-Hall, Inc. New Jersey, pp.247.
Pough, F.H., Janis, C.M., and Heiser, J.B. 2005. Vertubrate Life, 7th ed. Pearson Education, Inc. New Jersey, pp.74.
Rasmussen, A.R. 1997. Systematics of sea snakes: a critical review. Symp. Zool. Soc. Lond.(70):15-30.
Ruben, J.A. 1977. Morphological correlates of predatory modes in the coachwhip (Masticophis flagellum) and rosy boa(Lichanura roseofusca). Herpetologica.(33):1-6.
Saint Girons, H. 1964. Notes sur l'ecologie et la structure des populations Des Laticaudinae(Serpentes, Hydrophiidae) en Nouvelle Caledonie. Terre. la. Vie.(111):185-214.
Scanlon, J.D., and Lee, M.S.Y. 2004. Phylogeny of Australasian venomous snakes(Colubridea, Elapidae, Hydrophiinae) based on phenotypic and molecular evidence. Zool. Scr.(33):335-366.
Schulte-Hostedde, A.I., Zinner, B., Sillar, B.Z., and Hickling, G.J. 2005. Restitution of mass-size residuals: Validating body condition indices. Ecology(85)1:155-163.
Seigel, J.T. Collins, and S.S. Novak(eds.), Macmillan, Inc. New York, pp.442-477.
Seymour, R.S. 2000. Model analogues in the study of cephalic circulation. Comp. Biochem. Physiol. A.(125):517-524.
Shetty, S., ane Prasad, K.V.D. 1996. Studies on the terrestrial behaviour of Laticauda colubrina in the Andaman Islands, India. Hamadryad(21):23-26.
Shine, R., Cogger, H.G., Reed, R.R., Shetty, S. and Bonnet, X. 2003. Aquatic and torrestrial locomotor speeds of amphibious sea-snakes(Serpentes, Laticaudidae). J. Zool., Lond.(259):261-268.
Shine, R., and Shetty, S. 2001, Moving in two worlds - aquatic and terrestrial locomotion in sea snakes(Laticauda colubrina, Laticaudae), J. Evol. Biol.(14):338-346.
Slowinski, J.B., Knight, A., and Rooney, A.P. 1997. Inferring species trees from gene trees: A phylogenetic analysis of the Elapidae (Serpentes) based on amino acid sequences of neurotoxins. Mol. Phylogenet. Evol.(8):349-362.
Smith, H.M., Smith, R.B., and Sawin, H.L. 1977. A summary of snake classification (Reptilia, Serpentes). J. Herpetol.(11):125-121.
Srygley, R.B., and Kingsolver, J.G. 2000. Effects of weight loading on flight performance and survival of palatable neotropical Anartia fatima butterflies. Biol. J. Linn. Soc.(70):707-725.
Stewart, G.R. 1965. Thermal ecology of the garter snakes Thamnophis Sirtalis concinnus(Hallowell) and Thamnophis ordinoides(Baird and Girard). Herpetologica(21):81-103.
Stewart, J.R. 1984. Thermal biology of the live bearing lizard Gerrhonotus coeruleus. Herpetologica(40):349-355.
Stuebing, R.B. 1988. Island romance: the biology of the yellow-lipped sea krait. Malay. Nat.(41):9-11.
Vanhooydonck, B. and Van Damme, R. 2001. Evolutionary trade-offs in locomotor capacities in lacertid lizards: are splendid sprinters clumsy climbers?. J . Evol. Biol.(14):46-54.
Van Damme, R., Aerts. P., and Vanhooydonck, B. 1997. No trade-off Between sprinting and climbing in two populations of the Lizard Podarcis hispanica(Reptilia: Lacertifae). Biol. J. Linn. Soc.(60):493-503.
Voris, H.K. 1975. Dermal scale-vertebra relationships in sea snakes (Hydrophiiade). Copeia(4):746-757.
Voris, K.H. and Jayne, C.B. 1976. The costocutaneous muscles in some sea snakes(Reptilia, Serpentes). J. Herpetol.(10):175-180.
Wiefemann, E. 1932. Zur ortsbewegung der Schlangen und Schleichen. Zool. Jahrb. Abt. Allg. Zool. Physiol. Tiere.(50):557-596.
太田英利，增永元。2005。琉球のウミヘビ—その多様性と自然史。「南の島の自然誌—沖縄と小笠原の海洋生物研究のフィールドから」。矢野和成。東海大学出版会。日本。神奈川県。159-172頁。
毛壽先，陳本源。1981。台灣海蛇之研究。國防部軍醫局印製。台灣，台北。62頁。
杜銘章。1987。蘭嶼闊尾青斑海蛇(Laticauda semifasciata)生態之研究。國立中山大學海洋生物研究所碩士論文。台灣。高雄。
杜銘章。2004。蛇類大驚奇。遠流出版事業股份有限公司。台灣。台北。
杜銘章，林郁君，黃基礎。1999。影響蛇類跨越能力的變因。師大生物學報(34)2:129-135.
呂光洋，杜銘章，向高世。1999。台灣兩棲爬蟲圖鑑。大自然雜誌社。台灣。台北。
林登秋。2007。分布的形態。「生物地理學—從生態與演化的角度探討」。鄭勝華，林登秋，徐堉峰，呂光洋（主編）。藝軒圖書出版社。台灣，台北。73-117頁。
張鈞睿。2010。台灣三種闊尾海蛇地理分佈及陸棲程度與溫度耐受及偏好之關係。國立台灣師範大學生命科學系碩士論文。台灣。台北。
森哲。1994。ウミヘビ学入門ー海で生きるヘビ達。みとりいし。(5):23-26。
黃雯杰。2008。眼鏡蛇(Naja atra)的血壓調節研究。國立台灣師範大學生命科學系碩士論文。台灣。台北。